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So basically, a RTX 4090 (PCIe 4) installed in a PCIe 5 slot will run at PCIe 4 speeds. If the slot is an x8 slot, it will run at PCIe 4 x8 - which is the same as PCIe 3 x16 in terms of speed.
2 RAM Slots Motherboard | Pangoly
The M.2 and x16 PCIe slots on a motherboard house the Ryzen 5 3600 must run PCIe 4.0. With this card, a reasonable amount of RAM clock speed overclocking should be possible.
TS-877XU-RP-3600-8G
Slot 1: PCIe 3.0 x16 or PCIe 3.0 x8; full-height, full-length (PCIe x16 slot is double-wide) · Slot 2: PCIe 3.0 x8; full-height, full-length (not present if the ...